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Prof. Ryland, John

University of Wales Swansea; Biological Sciences, more

Function: Emeritus Professor of Marine Biology
Direct contact at institute:
Tel.: +44-(0)1792-29 54 40
E-mail:

A1 Publications (11) [show]
Ryland, J.S.; Bishop, J.D.D.; De Blauwe, H.; El Nagar, E.; Minchin, D.; Wood, C.A.; Yunnie, A.L.E. (2011). Alien species of Bugula (Bryozoa) along the Atlantic coasts of Europe. Aquat. Invasions 6(1): 17-31. https://dx.doi.org/10.3391/ai.2011.6.1.03, more Ryland, J. (2009). John Gray (1941-2007): a tribute. Mar. Ecol. (Berl.) 30(s1): 200-202. https://dx.doi.org/10.1111/j.1439-0485.2009.00310.x, more Ryland, J.S. (2009). Book review: Mosdiertjes van de Zuidelijke bocht van de Noordzee [Moss animals of the Southern Bight of the North Sea] by Hans De Blauwe, Ostende, Belgium, Vlaams Instituut voor de Zee (VLIZ), 2009, 464 pp. ISBN 978-90-812-9003-6. J. Nat. Hist. 43(41-44): 2787-2788, more Ryland, J.S.; Lord, R.; Mackie, J.A. (2009). Recent discoveries of alien Watersipora (Bryozoa) in Western Europe, with redescriptions of species. Zootaxa 2093(2093): 43-59, more Ryland, J.S.; Porter, J.S. (2003). The identity of Alcyonidium gelatinosum (Linnaeus, 1761) (Bryozoa: Ctenostomatida). J. Nat. Hist. 37: 2179-2189. https://dx.doi.org/10.1080/00222930210138917, more Ryland, J.S.; de Putron, S.; Scheltema, R.S.; Chimonides, P.J.; Zhadan, D.G. (2000). Semper's (zoanthid) larvae: pelagic life, parentage and other problems. Hydrobiologia 440(1-3): 191-198. https://dx.doi.org/10.1023/A:1004171127777, more Ryland, J.S.; Porter, J.S. (2000). Alcyonidium reticulum sp. nov., a common intertidal bryozoan from south-west Britain. J. Mar. Biol. Ass. U.K. 80(3): 563-564. https://dx.doi.org/10.1017/S0025315400002344, more Ryland, J.S.; Bishop, J.D.D. (1993). Internal fertilisation in hermaphroditic colonial invertebrates, in: Ansell, A.D. et al. Oceanogr. Mar. Biol. Ann. Rev. 31. Oceanography and Marine Biology: An Annual Review, 31: pp. 445-477, more Ryland, J.S. (1976). Physiology and ecology of marine bryozoans, in: Russell, F.S. et al. Adv. Mar. Biol. 14. Advances in Marine Biology, 14: pp. 285-443, more Ryland, J.S. (1967). Polyzoa, in: Barnes, H.B. (Ed.) Oceanogr. Mar. Biol. Ann. Rev. 5. Oceanography and Marine Biology: An Annual Review, 5: pp. 343-369, more Ryland, J.S. (1958). Bugula simplex Hincks, a newly recognized polyzoan from British waters. Nature (Lond.) 181: 1146-1147. https://dx.doi.org/10.1038/1811146b0, more
Peer reviewed publications (3) [show]
Hayward, P.J.; Ryland, J.S. (1991). New and little-known Bryozoa from Antarctica and the southwest Atlantic. Bull. Mus. Natl. Hist. Nat., Sect. A Zool. Biol. Ecol. Anim. Section A, 13(3-4): 241-261, more Ryland, J.S. (1974). A revised key for the identification of intertidal bryozoa (Polyzoa). Field Stud. 4(1): 77-86, more Ryland, J.S. (1960). The British species of Bugula (Polyzoa). Proc. Zool. Soc. Lond. 134: 65-105, more
Books (14) [show]
Hayward, P.J.; Ryland, J.S. (Ed.) (2017). Handbook of the marine fauna of North-West Europe. Second edition. Oxford University Press: Oxford. ISBN 978-0-19-954944-3. xiii, 785 pp., more Relini, G.; Ryland, J. (Ed.) (2007). Biodiversity in enclosed seas and artificial marine habitats: Proceedings of the 39th European Marine Biology Symposium, held in Genoa, Italy, 21-24 July 2004. European Marine Biology Symposia, 39. Developments in Hydrobiology, 193. ISBN 978-1-4020-6155-4; e-ISBN 978-1-4020-6156-1. VII, 271 pp. https://dx.doi.org/10.1007/978-1-4020-6156-1, more Hayward, P.J.; Ryland, J.S. (1999). Cheilostomatous Bryozoa: 2. Hippothooidea - Celleporoidea: notes for the identification of British species. 2nd ed. Synopses of the British Fauna, N.S. 10. Field Studies Council: Shrewsbury. ISBN 1-85153-263-3. VII, 416 pp., more Hayward, P.J.; Ryland, J.S. (1998). Cheilostomatous Bryozoa: 1. Aeteoidea - Cribrilinoidea: notes for the identification of British species. 2nd ed. Synopses of the British Fauna, N.S. 10. Field Studies Council: Shrewsbury. ISBN 1-85153-261-7. VII, 366 pp., more Hayward, P.J.; Ryland, J.S. (Ed.) (1995). Handbook of the marine fauna of North-West Europe. First edition. Oxford University Press: Oxford. ISBN 0-19-854054-X. XI, 800 pp., more Hayward, P.J.; Ryland, J.S.; Taylor, P.D. (Ed.) (1994). Biology and palaeobiology of bryozoans: Proceedings of the 9th International Bryozoology Conference, School of Biological Sciences, University of Wales, Swansea, 1992. International Symposium Series, 9. Olsen & Olsen: Fredensborg. ISBN 87-85215-23-6. VIII, 240 pp., more Hayward, P.J.; Ryland, J.S. (Ed.) (1990). The marine fauna of the British Isles and North-West Europe: 1. Introduction and protozoans to arthropods. Clarendon Press: Oxford. ISBN 0-19-857356-1. xvi, 1-627 pp., more Hayward, P.J.; Ryland, J.S. (Ed.) (1990). The marine fauna of the British Isles and North-West Europe: 2. Molluscs to chordates. Clarendon Press: Oxford. ISBN 0-19-857515-7. xvi, 628-996, pp., more Ryland, J.S.; Tyler, P.A. (Ed.) (1989). Reproduction, genetics and distributions of marine organisms: 23rd European Marine Biology Symposium, School of Biological Sciences, University of Wales, Swansea, 5-9 September 1988. European Marine Biology Symposia, 23. Olsen & Olsen: Fredensborg. ISBN 87-85215-15-5. VIII, 469 pp., more Hayward, P.J.; Ryland, J.S. (1985). Cyclostome bryozoans: keys and notes for the identification of the species. Synopses of the British Fauna, N.S. 34. E.J. Brill/W. Backhuys: London. ISBN 90-04-07697-2. VI, 154 pp., more Hayward, P.J.; Ryland, J.S. (1979). British ascophoran bryozoans. Keys and notes for the identification of the species. Synopses of the British Fauna, N.S. 14. Academic Press: London, New York and San Francisco. ISBN 0-12-335050-6. 312 pp., more Ryland, J.S.; Hayward, P.J. (1977). British anascan bryozoans. Cheilostomata: Anasca: keys and notes for the identification of the species. Synopses of the British Fauna, N.S. 10. Academic Press: London. ISBN 0-12-605250-6. 118 pp., more Ryland, J.S. (1965). Catalogue des principales sallissures marines (rencontrées sur les coques de navires dans les eaux européennes). Catalogue des principales salissures marines (rencontrées sur les coques de navires dans les eaux européennes), 2. Organisation de Coopération et de Développement Economiques: Paris. 82 pp., more Ryland, J.S. (1965). Catalogue of main marine fouling organisms. Found on ships coming into European waters. Volume 2: Polyzoa. OECD Publishing: [s.l.]. 83 pp., more
Book chapters (17) [show]
Cornelius, P.F.S.; Manuel, R.L.; Ryland, J.S.; Wood, C. (2017). Hydroids, sea anemones, jellyfish, and comb jellies: Phylum Cnidaria, in: Hayward, P.J. et al. (Ed.) Handbook of the marine fauna of North-West Europe. pp. 65-132, more Knight-Jones, E.W.; Ryland, J.S. (2017). Priapulids, sipunculans, echiurans, and entoprocts: Phylum Cephalorhyncha, in: Hayward, P.J. et al. (Ed.) Handbook of the marine fauna of North-West Europe. pp. 271-281, more Knight-Jones, E.W.; Ryland, J.S. (2017). Acorn worms and sea squirts: Phylum Hemichordata, in: Hayward, P.J. et al. (Ed.) Handbook of the marine fauna of North-West Europe. pp. 665-691, more Ryland, J.S. (2017). Bryozoa, in: Hayward, P.J. et al. (Ed.) Handbook of the marine fauna of North-West Europe. pp. 603-637, more Ryland, J.S.; Lancaster, J. (2017). Fish, in: Hayward, P.J. et al. (Ed.) Handbook of the marine fauna of North-West Europe. pp. 693-740, more Ryland, J.S.; Porter, J.S. (2013). Species of Alcyonidium (Ctenostomatida) from the Pacific coast of North America: A preliminary account, in: Ernst, A. et al. Bryozoan Studies 2010. Lecture Notes in Earth System Sciences, 143: pp. 289-302. http://dx.doi.org/10.1007/978-3-642-16411-8_19, more Ryland, J.S.; de Putron, S.; Scheltema, R.S.; Chimonides, P.J.; Zhadan, D.G. (2000). Semper's (zoanthid) larvae: pelagic life, parentage and other problems, in: Jones, M.B. et al. Island, Ocean and Deep-Sea Biology: Proceedings of the 34th European Marine Biology Symposium, held in Ponta Delgada (Azores), Portugal, 13-17 September 1999. Developments in Hydrobiology, 152: pp. 191-198. https://dx.doi.org/10.1007/978-94-017-1982-7_18, more Cadman, P.S.; Ryland, J.S. (1996). The characters, reproduction, and growth of Alcyonidium mytili Dalyell, 1848 (Ctenostomatida), in: Gordon, D.P. et al. Bryozoans in space and time: proceedings of the 10th International Bryozoology Conference, Victoria University of Wellington, Wellington, New Zealand, 1995. pp. 69-79, more Cornelius, P.F.S.; Manuel, R.L.; Ryland, J.S. (1996). Hydroids, sea anemones, jellyfish and comb jellies ( Phylla Cnidaria and Ctenophora), in: Hayward, P.J. et al. (Ed.) (1995). Handbook of the marine fauna of North-West Europe. pp. 62-135, more Knight-Jones, E.W.; Ryland, J.S. (1996). Priapulids, sipunculans, echiurans and entoprocts (Phyla Priapulida, Sipuncula, Echiura and Entoprocta), in: Hayward, P.J. et al. (1995). Handbook of the marine fauna of North-West Europe. pp. 278-288, more Knight-Jones, E.W.; Ryland, J.S. (1996). Acorn-worms and sea squirts (Phyla Hemichordata and Urochordata), in: Hayward, P.J. et al. (Ed.) (1995). Handbook of the marine fauna of North-West Europe. pp. 687-711, more Ryland, J.S. (1996). Fish (Subphylum Euchordata (Vertebrata) Pisces), in: Hayward, P.J. et al. (Ed.) (1995). Handbook of the marine fauna of North-West Europe. pp. 712-757, more Ryland, J.S. (1996). Bryozoans (Phylum Bryozoa), in: Hayward, P.J. et al. (Ed.) (1995). Handbook of the marine fauna of North-West Europe. pp. 629-661, more Yonow, N.; Ryland, J.S. (1992). Growth and life history parameters in Acteon tornatilis (L.) (Opisthobranchia: Cephalaspidea), in: Colombo, G. et al. (Ed.) Marine Eutrophication and Population Dynamics: 25th European Marine Biology Symposium, Ferrara (Italy), 10-15 September 1990. International Symposium Series, : pp. 271-276, more Hayward, P.J.; Ryland, J.S. (1975). Growth, reproduction and larval dispersal in Alcyonidium hirsutum (Fleming) and some other Bryozoa, in: Bonaduce, G. et al. (Ed.) 8th European Marine Biology Symposium Sorrento (Naples) 1973. Pubblicazioni della Stazione Zoologica di Napoli, 39(Suppl. 1): pp. 226-241, more Ryland, J.S. (1975). Parameters of the lophophore in relation to population structure in a bryozoan community, in: Barnes, H.B. (Ed.) Ninth European Marine Biology Symposium. pp. 363-393, more Ryland, J.S.; Stebbing, A.R.D. (1971). Settlement and orientated growth in epiphytic and epizoic bryozoans, in: Crisp, D.J. (Ed.) Fourth European Marine Biology Symposium. pp. 105-123, more
Report [show]
Ryland, J.S.; Hayward, P.J. (1991). Marine flora and fauna of the Northeastern United States: erect Bryozoa. NOAA Technical Report, NMFS(99). U.S. Department of Commerce. NOAA. National Marine Fisheries Service: Washington D.C. 47 pp., more
Other publications (3) [show]
Ryland, J.S. (1966). Observations on the development of larvae of the Plaice, Pleuronectes platessa L., in aquaria. J. Cons. - Cons. Int. Explor. Mer 30(2): 177-195, more Ryland, J.S. (1965). Polyzoa (Bryozoa). Order: Cheilostomata, Cyphonautes Larvae. Fiches d'identification du zooplancton, 107. ICES: Copenhagen. 6 pp., more Ryland, J.S. (1962). Some species of Bugula (Polyzoa) from the Bay of Naples. Pubbl. Stn. Zool. Napoli 33: 20-31, more

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